1International Joint Research Laboratory for Global Change Ecology, School of Life Sciences, Henan University, Kaifeng 475004, China; 2Henan Dabieshan National Field Observation and Research Station of Forest Ecosystem, Zhengzhou 450046, China; 3Key Laboratory of Geospatial Technology for the Middle and Lower Yellow River Regions, Ministry of Education, College of Environment and Planning, Henan University, Kaifeng 475004, China; 4Henan Key Laboratory of Integrated Air Pollution Control and Ecological Security, Kaifeng 475004, China; 5Department of Ecology and Evolutionary Biology, University of Michigan, Michigan 48109, USA; 6South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China; 7Department of Natural Resource of Henan Province, Institute of Natural Resources Survey and Planning, Zhengzhou 450000, China