1Faculty of Life and Environmental Sciences, University of Tsukuba, Tsukuba, Japan; 2Department of Biology, School of Arts and Sciences, National University of Mongolia, Ulaanbaatar, Mongolia; 3Botanic Garden and Research Institute, Mongolian Academy of Science, Ulaanbaatar, Mongolia; 4Graduate School of Environmental Studies, Nagoya University, Nagoya, Japan; 5Arid Land Research Center, Tottori University, Tottori, Japan