1 Department of Ecology and Ecological Engineering, College of Resources and Environmental Sciences, and Key Laboratory of
Biodiversity and Organic Farming of Beijing City, China Agricultural University, Beijing 100193, China, 2 Institute of Ecology and Key
Laboratory for Earth Surface Processes of the Ministry of Education, College of Urban and Environmental Sciences, Peking University,
Beijing 100871, China, 3 State Key Laboratory Breeding Base of Dao-di Herbs, National Resource Center for Chinese Materia Medica,
China Academy of Chinese Medical Science, Beijing 100700, China