1Zaozhuang University, Zaozhuang 277000, China, 2Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China, 3State Key Laboratory of Tree Genetics and Breeding, Institute of Ecological Conservation and Restoration, Chinese Academy of Forestry, Beijing 100091, China, 4Gansu Dunhuang-Kumtag Desert Ecosystem Observation and Research Station, Dunhuang 736200, China, 5Experimental Center of Desert Forestry, Chinese Academy of Forestry, Dengkou 015200, China, 6Kunming Institute of Botany, Chinese Academy of Sciences, Kunming 650201, China, 7Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan 430074, China, 8Institute of Great Green Wall, Bayannur 015200, China, 9Institute of Desertifcation Studies, Chinese Academy of Forestry, Beijing 100091, China
This study was fnancially supported by the Shandong Provincial Natural Science Foundation (ZR2022QC259), National Natural Science Foundation of China (41571270) and China Postdoctoral Science Foundation (2024M753463).