1CAS Key Laboratory of Aquatic Botany and Watershed Ecology, Wuhan Botanical Garden, Chinese Academy of Sciences, Wuhan
430074, Hubei, China, 2 University of Chinese Academy of Sciences, Beijing 100049, China, 3 Core Botanical Garden, Chinese Academy
of Sciences, Wuhan 430074, Hubei, China, 4 School of Life Sciences, Henan University, Kaifeng 475004, Henan, China